BowDown
Spends too much time on here
Shift forks and clutches develop wear ridges over time running in the same position. Motor commutators will oxidize. Gear and cam lubes will drain off due to gravity. It's a good idea to operate things once in awhile to redistribute all that stuff, and also verifies that it actually works. And if not, clues you in to visit the shop for repair, rather than be surprised when you needed it.
I also like to run the engine at high RPM once in awhile. Wear ridges build up near the tops of cylinders from the piston rings, and the connecting rods stretch and compress a bit more with RPM. So if I were to need full power in an emergency, I don't want it to be the first time the rings have encountered that particular bit of cylinder wall.
There's no redline on the tachometer, so it must be good at any speed up to 8,000, right?
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All true. I have a friend that had a nissan titan 4x4. He rarely used 4x4 and tried on day when he needed it, system wouldn't engage. Took it to nissan and they had to replace the entire front axle (diff not serviceable alone) due to no lubrication from none use